Beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died at about 141, the zoo says, adding that "caring for such a remarkable tortoise was a privilege." ...

One Galápagos tortoise lived 141 years, turning quiet meals and slow steps into a legacy of care, memory, and conservation ...

The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the ...

An occasional series where Mongabay founder Rhett Ayers Butler shares analysis, perspectives and story summaries. She moved ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Giant tortoises are some of the long-lived species on the planet. Here’s what biologists now understand about one of nature’s ...

The beloved Galapagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has died LOS ANGELES -- After more than a century of munching on her favorite ...

LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.