The California zoo invited fans of the late Galápagos tortoise to enjoy one of her favorite snacks, a "generous, fruit-filled salad," in her honor ...

One Galápagos tortoise lived 141 years, turning quiet meals and slow steps into a legacy of care, memory, and conservation ...

The beloved Galápagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has died. Zoo officials say she died on Nov. 20.

The San Diego Zoo announced Gramma, the Galápagos tortoise, died on Nov. 20. She was about 141 years old based on estimates from experts.

Gramma, who died at the zoo on Thursday, was believed to have been born in 1884, when Chester A. Arthur was in the White House.

Gramma was described as "a quiet and constant presence at the zoo, a witness to history, a beloved icon, and an extraordinary ambassador for her species." ...

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the Oval Office and there were only 39 states at the time.

San Diego Zoo mourns the loss of Gramma, a Galápagos tortoise estimated to be 141 years old. Born in the Galápagos Islands, ...

Gramma, one of San Diego Zoo’s most beloved residents, has died at around 141 years old, zoo officials said. She was born in ...

Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died at an estimated 141 years of ...

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.